Adrian, Janick and Dave's guitars and Steve's basses

Not during the solo. That's an octaver effect. Probably a Boss one or a Pog.
It would most likely be an effect that's provided by the Roland GR700 guitar synth. The Synth Controller (ie the guitar, or the midi pickup mounted on the guitar and its associated control knobs) was the Lado Lazer, which had the MIDI pickup and control knobs from the Roland G-707 installed/mounted on it (Just like the Jackson he used on the previous tour).
 
It would most likely be an effect that's provided by the Roland GR700 guitar synth. The Synth Controller (ie the guitar, or the midi pickup mounted on the guitar and its associated control knobs) was the Lado Lazer, which had the MIDI pickup and control knobs from the Roland G-707 installed/mounted on it (Just like the Jackson he used on the previous tour).
I mean the versions from 2003 onwards.

1988 was the Roland. The lower octave.
 
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